Centre for Ecological Research

Institute of Ecology and Botany

"Lendület" Landscape and Conservation Ecology

Publications of Attila Torma

pdf: link to open access pdf at the journal's homepage.

abstract: link to abstract at the journal's homepage. If you need a copy of any of these publications, please, drop me an e-mail.


  • Šeat, J. & Torma, A. 2021. How steady is the nested pattern in saline grassland true bug communities? Effects of sampling effort and data completeness on nestedness. - Acta Oecologica 110: 103670. pdf


  • Maák, I., Tóth, E., Lenda, M., Lőrinczi, G., Kiss, A., Juhász, O., Czechowski, W. & Torma, A. 2020. Behaviours indicating cannibalistic necrophagy in ants are modulated by the perception of pathogen infection level. - Scientific Reports 10: 17906. pdf

  • Deák, B., Valkó, O., Nagy, D.D., Török, P., Torma, A., Lőrinczi, G., Kelemen, A., Nagy, A., Bede, Á., Mizser, Sz., Csathó, A.I. & Tóthmérész, B. 2020. Habitat islands outside nature reserves – Threatened biodiversity hotspots of grassland specialist plant and arthropod species. - Biological Conservation 241: 108254. pdf


  • Kaur, H., Torma, A., Gallé-Szpisjak, N., Šeat, J., Lőrinczi, G., Módra, G. & Gallé, R. 2019. Road verges are important secondary habitats for grassland arthropods. - Journal of Insect Conservation 23: 899–907. pdf

  • Torma, A., Császár, P., Bozsó, M., Deák, B. Valkó, O. Kiss, O. & Gallé, R. 2019. Species and functional diversity of arthropod assemblages (Araneae, Carabidae, Heteroptera and Orthoptera) in grazed and mown salt grasslands. - Agriculture, Ecosystems & Environment 273: 70–79. abstract


  • Gallé, R., Császár, P., Makra, T., Gallé-Szpisjak, N., Ladányi, Z., Torma, A., Ingle, K. & Szilassi, P. 2018. Small-scale agricultural landscapes promote spider and ground beetle densities by offering suitable overwintering sites. - Landscape Ecology 33: 1435–1446. pdf

  • Gallé, R., Szabó, Á., Császár, P. & Torma, A. 2018. Spider assemblage structure and functional diversity patterns of natural forest steppes and exotic forest plantations. - Forest Ecology and Management 411: 234–239. abstract

  • Császár, P., Torma, A., Gallé-Szpisjak, N., Tölgyesi, Cs. & Gallé, R. 2018. Efficiency of pitfall traps with funnels and/or roofs in capturing ground-dwelling arthropods. - European Journal of Entomology 115: 15–24. pdf

  • Tölgyesi, C., Császár, P., Torma, A., Török, P., Bátori, Z. & Gallé, R. 2018. Think twice before using narrow buffers: Attenuating mowing-induced arthropod spillover at forest–grassland edges. - Agriculture, Ecosystems & Environment 255: 37–44. abstract

  • Torma, A., Bozsó, M. & Gallé, R. 2018. Secondary habitats are important in biodiversity conservation: a case study on orthopterans along ditch banks. - Animal biodiversity and conservation 41: 97–108. pdf


  • Torma, A., Bozsó, M., Tölgyesi, C. & Gallé, R. 2017. Relationship of different feeding groups of true bugs (Hemiptera: Heteroptera) with habitat and landscape features in Pannonic salt grasslands. - Journal of Insect Conservation 21: 645–656. pdf

  • Gallé, R., Gallé-Szpisjak, N. & Torma, A. 2017. Habitat structure influences the spider fauna of short-rotation poplar plantations more than forest age. - European Journal of Forest Research 131: 51–58. pdf


  • Gallé, R., Szpisjak, N. & Torma, A. 2016. Influence of habitat structure on the spiders of river islands and floodplain forests of the lower reach of the Mures River in Western Romania. - North-Western Journal of Zoology 12: 255–260. pdf

  • Maák, I., Torma, A., Kovács, J., Somogyi, A. & Lőrinczi, G. 2016. Threat, signal or waste? Meaning of corpses in two dulotic ant species. - Journal of Insect Behavior 29: 432–448. pdf

  • Gallé, R., Torma, A. & Maák, I. 2016. The effect of forest age and habitat structure on the ground-dwelling ant assemblages of lowland poplar plantations. - Agricultural and Forest Entomology 18: 151–156. abstract

  • Bátori, Z., Körmöczi, L., Zalatnai, M., Erdős, L., Ódor, P., Tölgyesi, C., Margóczi, K., Torma, A., Gallé, R., Cseh, V. & Török, P. 2016. River dikes in agricultural landscapes: The importance of secondary habitats in maintaining landscape-scale diversity. - Wetlands 36: 251–264. pdf

  • Torma A. & Bozsó M. 2016. Effects of habitat and landscape features on grassland Orthoptera on floodplains in the lower reaches of the Tisza River Basin. - European Journal of Entomology 113: 60–69. pdf


  • Torma, A., Gallé, R. & Bozsó, M. (2014). Effects of habitat and landscape characteristics on the arthropod assemblages (Araneae, Orthoptera, Heteroptera) of sand grassland remnants in Southern Hungary. - Agriculture, Ecosystems & Environment 196: 42–50. abstract


  • Torma, A. & Császár, P. 2013. Species richness and composition patterns across trophic levels of true bugs (Heteroptera) in the agricultural landscape of the lower reach of the Tisza River Basin. - Journal of Insect Conservation 17: 35–51. pdf


  • Torma, A. & Rédei, D. 2012. Additions and corrections to the checklist of true bugs of Hungary (Hemiptera: Heteroptera).- Natura Somogyiensis 22: 53–56. pdf

  • Gallé, R., Lőrinczi, G., Szpisjak, N., Maák, I. & Torma, A. (2012): Data on the arthropod (Araneae, Formicidae, Heteroptera) fauna of floodplain forests at the lower reach of the river Maros/Mures. In: Körmöczi, L. (ed.) Landscape-scale connections between the land use, habitat quality and ecosystem goods and services in the Mures/Maros valley. - Tiscia Monograph Series 10: 45–66. pdf


  • Torma, A. & Gallé, R. 2011. Fine scale pattern of true bug assemblages (Heteroptera) across two natural edges. - Acta Zoologica Academiae Scientiarum Hungaricae 57: 367–383. pdf

  • Lőrinczi, G., Bozsó, M., Duma, I., Petrescu, M., Gallé, R. & Torma, A. 2011. Preliminary results on the invertebrate fauna (Araneae, Orthoptera, Heteroptera and Hymenoptera: Formicidae) of alkaline grasslands of the Hungarian-Romanian border. In: Körmöczi, L. (ed.) Ecological and socio-economic relations in the valleys of river Körös/Cris and river Maros/Mures. - Tiscia Monograph Series 9: 159–173. pdf


  • Gallé, R., Torma, A. & Körmöczi, L. 2010. Small scale effect of habitat heterogeneity on invertebrate assemblages in Hungarian sandy grasslands. - Polish Journal of Ecology 58: 333–346. abstract

  • Torma, A., Varga, Cs., Varga, M. 2010. Spatial pattern of true bugs (Heteroptera) in heterogeneous grassland – Preliminary results. - Acta Phytopathologica et Entomologica Hungarica 45: 81–87. abstract


  • Torma, A. & Körmöczi, L. 2009. The influence of habitat heterogeneity on the fine-scale pattern of an Heteroptera assemblage in a sand grassland. - Community Ecology 19, Suppl. 1: 413–425. pdf

  • Gallé, R. & Torma, A. 2009. Epigeic spider (Araneae) assemblages of natural forest edges in the Kiskunság (Hungary). - Community Ecology 10: 146–151. pdf

  • Torma, A. 2009. Data to the terrestrial Heteroptera fauna of Moldva. - Acta Scientiarum Transylvanica 17: 107–118. pdf

  • Torma, A. 2009. Data to the terrestrial Heteroptera fauna of Dobrogea. - Acta Scientiarum Transylvanica 17: 89–103. pdf


  • Torma, A. 2008. Contribution to the terrestrial heteroptera fauna of Northern-Dobrogea. - Scientific Annals of the Danube Delta Institute 14: paper 12.


  • Gallé, R., Torma, A. & Körmöczi, L. 2007. Composition of assemblages of ants (Hymenoptera: Formicidae) and associated arthropods (Heteroptera, Orthoptera, Araneae) at natural forest edges – Preliminary results. - In: Tajovský, K., Schlaghamerrský, J., Pižl, V. (eds.)- Contributions to Soil Zoology in Central Europe II. České Budějovize pp. 47–52.

  • Árendás, O., Bozsó, M., Gallé, R., Torma, A. & Körmöczi, L. 2007. Composition of assemblages of ants (Hymenoptera: Formicidae) and associated arthropods (Heteroptera, Orthoptera, Araneae) at natural forest edges.- Myrmecological News 10: 100. pdf


  • Torma, A. 2005. Three new and a rare true bug species in the Hungarian fauna (Hetroptera: Dipsocoridae, Reduviidae, Lygaeidae). - Folia Entomologica Hungarica 66: 35–38. pdf


  • Rédei, D. & Torma, A. 2003. Occurrence of the Southern Green Stink Bug, Nezera viridula (Heteroptera: Pentatomidae) in Hungary - Acta Phytopathologica et Entomologica 38: 365–367. abstract